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On deep roots of Iranian hunter-gatherers, and possible scenarios on their formation

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The Iranian hunter-gatherers are among the more enigmatic lineages, with unclarified deeper roots. They fall broadly within the West Eurasian cluster, but are placed in an "extreme" position, displaying no strong genetic affilation with nearby Anatolian or Levant groups, pointing to deep divergence and or partial distinct ancestries. Their genetic makeup and formation remains unclear, but it has been agreed that they are primarily derived from a deep local, presumably Basal Eurasian source, and can be modeled via a four-way admixture, including a Caucasus_UP component, an Ancient North Eurasian (ANE) component, and Önge-like component, and an extra Basal Eurasian component (Amjadi et al. 2025, Allentoft et. al. 2024, Lazaridis 2018 etc.). Neolithic Iranians display the highest amount of Basal Eurasian ancestry of all sequenced populations. The inferred Basal Eurasian lineage is assumed to have split from other Eurasians after the OoA exit, but before the archaic introgressio...